Repo APT Security - apt-get アップデート GPG 署名が無効

aptlyを使用して新しいaptリポジトリを設定し、リポジトリに署名し、aptlyを使用してリポジトリを提供し、aptキーリングにパブリックgpgキーを追加した後、次のコマンドでエラーが発生しますapt update。

Err:3 http://#REPO_URL#/#NAME# #DISTRIBUTION# InRelease                                    
  The following signatures were invalid: #KEY_ID#
Hit:4 http://apt.postgresql.org/pub/repos/apt sid-pgdg InRelease                  
Reading package lists... Done
W: GPG error: http://#REPO_URL#/#NAME# #DISTRIBUTION# InRelease: The following signatures were invalid: #KEY_ID#
E: The repository 'http://#REPO_URL#/#NAME# #DISTRIBUTION# InRelease' is not signed.
N: Updating from such a repository can't be done securely, and is therefore disabled by default.
N: See apt-secure(8) manpage for repository creation and user configuration details.

しかし、署名とgpgキーは大丈夫です。 InReleaseファイルのgpg署名が正常に確認されました。

curl http://#REPO_URL#/InRelease | gpg --keyring /etc/apt/trusted.gpg --verify
gpg: Signature made Wed 11 Jan 2017 04:01:23 PM CET
gpg:                using RSA key #KEY_ID#
gpg: Good signature from "#DESCRIPTION_GPG_KEY#" [unknown]
gpg: WARNING: This key is not certified with a trusted signature!
gpg:          There is no indication that the signature belongs to the owner.
Primary key fingerprint: #GOOD_KEY_FINGERPRINT#

また、同様のコマンドを使用してReleaseファイルを確認しましたRelease.gpg。

その間に何がapt update起こるのかを調べようとします。strace -o /tmp/strace -ff apt updategrep

grep 'apt.*key' ./strace*
./strace.29829:execve("/usr/bin/apt-key", ["/usr/bin/apt-key", "--quiet", "--readonly", "verify", "--status-fd", "3", "/tmp/apt.sig.ORUwxh", "/tmp/apt.data.kKXyrN"], [/* 28 vars */]) = 0
./strace.29829:open("/usr/bin/apt-key", O_RDONLY)      = 4
./strace.29888:execve("/usr/bin/apt-key", ["/usr/bin/apt-key", "--quiet", "--readonly", "verify", "--status-fd", "3", "/tmp/apt.sig.utRWBD", "/tmp/apt.data.Fo1Lka"], [/* 28 vars */]) = 0
./strace.29888:open("/usr/bin/apt-key", O_RDONLY)      = 4
./strace.29947:execve("/usr/bin/apt-key", ["/usr/bin/apt-key", "--quiet", "--readonly", "verify", "--status-fd", "3", "/tmp/apt.sig.ug6xiV", "/tmp/apt.data.Yv4zFs"], [/* 28 vars */]) = 0
./strace.29947:open("/usr/bin/apt-key", O_RDONLY)      = 4
./strace.30006:execve("/usr/bin/apt-key", ["/usr/bin/apt-key", "--quiet", "--readonly", "verify", "--status-fd", "3", "/tmp/apt.sig.QSyrCg", "/tmp/apt.data.LK9DGO"], [/* 28 vars */]) = 0
./strace.30006:open("/usr/bin/apt-key", O_RDONLY)      = 4

このエラーをどのようにデバッグして修正できますか？